Emeritus Professor of Neurophysiology at the University of Turin. He was a researcher in Canberra and Chicago with the Nobel Prize John Eccles. He is Honorary Professor of Neurology at the Northwestern University of Chicago. He is a member of international Societies and Academies, like the Academia Europaea. National Academy of Sciences XL Medal in Physical and Natural Sciences and Premio Feltrinelli at the Accademia dei Lincei.